1. Section of Ecology, Behavior and Evolution, University of California San DiegoLa Jolla, CA 92093-0116, USA
2. Department of Paleobiology, National Museum of Natural History, Smithsonian InstitutionWashington, DC 20013-7012, USA
3. Department of Geophysical Sciences, University of Chicago5734 South Ellis Avenue, Chicago, IL 60637, USA
4. Department of Integrative Biology, University of California BerkeleyBerkeley, CA 94720, USA