1. UMR 5558 ‘Biométrie et Biologie Evolutive’, Université Claude Bernard–Lyon 1, 43, Boulevard du 11 Novembre 1918, 69622 Villeurbanne Cedex, France
2. Centre National d'Etudes et de Recherches Appliquées Cervidés–sanglier, BP 15 Gerstheim, F–67154 Erstein Cedex, France
3. Centre for Ecological and Evolutionary Synthesis (CEES), Department of Biology, University of Oslo, PO Box 1050 Blindern, N–0316 Oslo, Norway
4. The University Centre in Svalbard (UNIS), N–9170 Longyearbyen, Spitsbergen, Norway