1. Department of Ecology and Evolutionary Biology, University of California Santa Cruz, Santa Cruz, CA 95064, USA
2. Coastal Research Center, Marine Science Institute and Department of Ecology, Evolution, and Marine Biology, University of California Santa Barbara, Santa Barbara, CA 93106, USA
3. Division of Natural Sciences, Monterey Peninsula College, Monterey, CA 93940, USA
4. National Oceanic and Atmospheric Administration, National Marine Fisheries Service, Southwest Fisheries Science Center, Honolulu Laboratory, 2570 Dole Street, Honolulu, HI 96822, USA