1. Department of Ecology and Evolutionary Biology Graduate School of Life Sciences, University of TohokuAobayama, Sendai 980-8578, Japan
2. Marine Science Institute and Department of Ecology Evolution and Marine Biology, University of CaliforniaSanta Barbara, CA 93106, USA
3. Smithsonian Tropical Research InstituteApartado 0843-03092 Balboa, Ancon, Panama City, Panama