1. Centre for Population Biology, Silwood Park, Ascot, Berkshire SL5 7PY, UK
2. Unit of Parasitoid Systematics, CABI Bioscience UK Centre (Ascot), Department of Biology, Imperial College, Silwood Park, Ascot, Berkshire SL5 7PY, UK
3. Department of Entomology,The Natural History Museum, London SW7 5BD, UK