1. Centre of Excellence in Biological Interactions, Department of Biosciences, University of Helsinki, Viikinaari 1, Biocenter 3, PO Box 65, Helsinki 00014, Finland
2. University of Helsinki Tvärminne Zoological Station, Faculty of Biological and Environmental Sciences, J.A. Palménin tie 260, Hanko 10900, Finland
3. Centre of Excellence in Biological Interactions, Department of Biological and Environmental Sciences, University of Jyväskylä, Survontie 9, Jyväskylä 40014, Finland