1. Centre for Evolutionary Biology and Neuroecology Group, School of Animal Biology (M092), The University of Western Australia, Crawley, Perth, Western Australia 6009, Australia
2. Genetics, Ecology and Evolution Group, Institute of Integrative and Comparative Biology, University of Leeds, Miall Building, Leeds LS2 9JT, UK
3. School of Biological, Biomedical and Environmental Sciences, University of Hull, Kingston upon Hull HU6 7RX, UK