1. School of Biological Sciences and School of Earth Sciences, University of Bristol, Life Sciences Building, Bristol BS8 1TG, UK
2. Department of Biology, National University of Ireland, Maynooth, Co. Kildare, Ireland
3. Department of Life Science, The Natural History Museum, London SW7 5BD, UK
4. Institute of Biological, Environmental and Rural Sciences (IBERS), Aberystwyth University, Aberystwyth, Ceredigion SY23 3FG, UK
5. Faculty of Life Sciences, University of Manchester, Oxford Road, Manchester M13 9PL, UK