1. Department of Psychology, University of Lethbridge, Lethbridge, Alberta, Canada T1K 3M4
2. Applied Behavioural Ecology and Ecosystems Research Unit, UNISA, Johannesburg, South Africa
3. Institute of Biological and Environmental Sciences, University of Aberdeen, Aberdeen AB24 2TZ, UK
4. Marine Alliance Science and Technology for Scotland, University of Aberdeen, Aberdeen AB24 2TZ, UK