1. Biosciences and Living Systems Institute, University of Exeter, Stocker Road, Exeter EX4 4QD, UK
2. Department of Marine Biology and Oceanography, Institut de Ciències del Mar (CSIC), 08003 Barcelona, Spain
3. Earlham Institute, Norwich Research Park, Norwich, NR4 7UZ, UK
4. School of Biological Sciences, University of East Anglia, Norwich, NR4 7TU, UK