1. Department of Ecology and Evolutionary Biology, University of California Los Angeles, Los Angeles, CA 90095, USA
2. The Bartlett Faculty of the Built Environment, University College London, London WC1H 0QB, UK
3. Centre de Recherches sur la Cognition Animale, Centre de Biologie Intégrative, CNRS, Université de Toulouse, 31062 Toulouse, France
4. Department of Philosophy and the Institute for Simulation and Training, University of Central Florida, Orlando, FL 32826, USA