1. Department of Zoology, University of Oxford, South Parks Road, Oxford OX1 3PS, UK
2. Molecular Genetics Unit, Okinawa Institute of Science and Technology Graduate University, Onna, Okinawa 904-0495, Japan
3. Centro Andaluz de Biología del Desarrollo, Consejo Superior de Investigaciones Científicas/Universidad Pablo de Olavide, 41013 Sevilla, Spain
4. School of Biological Sciences, University of Essex, Colchester, Essex, UK