1. Research and Innovation Center, Fondazione Edmund Mach, San Michele all'Adige-38010, Italy
2. Randall Division of Cell and Molecular Biophysics, New Hunt's House, King's College London, London SE1 1UL, UK
3. Department of Life Science and Medical Bioscience, Waseda University, 1-104 Totsukamachi Shinjuku, Tokyo 169-8050, Japan
4. The Gurdon Institute, University of Cambridge, Tennis Court Road, Cambridge CB2 1QN, UK
5. Genetics Department, University of Cambridge, Downing Street, Cambridge CB2 3EH, UK