1. Atlas of Living Australia, CSIRO National Research Collections Australia, GPO Box 1700, Canberra ACT 2601, Australia
2. Land and Water, Commonwealth Scientific and Industrial Research Organisation (CSIRO), GPO Box 1600, Canberra ACT 2601, Australia
3. Centre for Biodiversity Analysis and Research School of Biology, The Australian National University, Acton ACT 2601, Australia