1. Department of Epidemiology and Biostatistics, College of Public Health, University of Georgia, Athens, GA 30602, USA
2. Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI 48109, USA
3. Center for the Study of Complex Systems, University of Michigan, Ann Arbor, MI 48109, USA
4. Fogarty International Center, National Institutes of Health, Bethesda, MD 20892, USA