1. School of Biological Sciences, University of East Anglia, Norwich Research Park, Norwich NR4 7TJ, UK
2. Department of Biology and CESAM—Centre for Environmental and Marine Studies, University of Aveiro, Campus de Santiago, 3810-193 Aveiro, Portugal
3. South Iceland Research Centre, University of Iceland, Lindarbraut 4, 840 Laugarvatn, Iceland