1. Laboratoire d'Ecologie Alpine, CNRS, Université Grenoble Alpes, Université Savoie Mont Blanc, 38000 Grenoble, France
2. Cirad, University of Montpellier, Montpellier 34098, France
3. Cifor, Lima 15024, Peru
4. Fenner School of Environment and Society, Australian National University, Linnaeus Way, Canberra, Australian Capital Territory 2601, Australia