1. Division of Biology, Imperial College London, Silwood Park Campus, Ascot, Berkshire SL5 7PY, UK
2. NERC Centre for Population Biology, Imperial College London, Silwood Park Campus, Ascot, Berkshire SL5 7PY, UK
3. Grantham Institute for Climate Change, Imperial College London, South Kensington Campus, London SW7 2AZ, UK