1. Department of Biology, University of York, York YO10 5DD, UK
2. Forest Research Centre, Sabah Forestry Department, PO Box 1407, 90715 Sandakan, Sabah, Malaysia
3. School of Sustainable Agriculture, Universiti Malaysia Sabah, 88400 Kota Kinabalu, Sabah, Malaysia
4. University of the Philippines Los Baños, Laguna, Philippines
5. Institute of Integrative and Comparative Biology, University of Leeds, Leeds LS2 9JT, UK