1. Porter Alliance, Centre for Environmental Policy, Imperial College London, London SW7 2AZ, UK
2. Natural Resources Management Centre, Department of Natural Resources, University of Cranfield, Bedford MK43 0AL, UK
3. Agri-Environment and Land Use Strategy, Food and Environment Research Agency, Sand Hutton, York YO41 1LZ, UK
4. Porter Alliance, Department of Biology, Imperial College London, London SW7 2AZ, UK