1. Department of Zoology, University of OxfordOxford OX1 3PS, UK
2. Department of Ecology, Evolution and Environmental Biology, Columbia UniversityNew York, NY 10027, USA
3. Sackler Institute for Comparative Genomics, American Museum of Natural HistoryNew York, NY 10024, USA
4. Warwick HRI, University of WarwickWarwick CV35 9EF, UK