1. Zoological Society of LondonRegent's Park, London NW1 4RY, UK
2. Department of Nature Conservation, Tshwane University of TechnologyPrivate Bag X680, Pretoria 0001, Republic of South Africa
3. The Tony and Lisette Lewis Foundation and Mammal Research Institute, University of PretoriaPretoria 0001, Republic of South Africa
4. Department of Ecology and Evolutionary Biology, University of California621 Charles E. Young Drive South, Los Angeles, Los Angeles, CA 90095-1606, USA