1. Department of Geology, Palaeontology and Mineralogy, Hessisches Landesmuseum Darmstadt, Friedensplatz 1, 64283 Darmstadt, Germany
2. Institute of Biomedical Life Sciences, Graham Kerr Building, University of Glasgow, Glasgow G12 8QQ, UK
3. Department of Entomology, San Diego Natural History Museum, PO Box 121390, San Diego, CA 92112, USA