1. Department of Biological Sciences, School of Life and Environmental Sciences, Royal Holloway University of London, Egham, Surrey TW20 0EX, UK
2. School of Engineering and Innovation, Open University, Milton Keynes MK7 6AA, UK
3. Department of Zoology, University of Cambridge, Downing Street, Cambridge CB2 3EJ, UK
4. Cambridge Institute of Therapeutic Immunology and Infectious Disease (CITIID), Jeffrey Cheah Biomedical Centre, Cambridge Biomedical Campus, Cambridge University, Cambridge CB2 0AW, UK
5. FitzPatrick Institute of African Ornithology, DST-NRF Centre of Excellence, University of Cape Town, Rondebosch 7701, Cape Town, South Africa
6. The Natural History Museum, Tring, Herts HP23 6AP, UK