1. Leiden University Centre for Linguistics, Leiden 2311 BE, The Netherlands
2. Copernicus Institute of Sustainable Development, Environmental Science Group, Utrecht University, Utrecht 3584 CB, The Netherlands
3. Department of Evolutionary Genetics, Max-Planck-Institute for Evolutionary Anthropology, Leipzig 04103, Germany
4. University of Groningen, Campus Fryslân 8911 CE, The Netherlands
5. Department of Anthropology, University of Wyoming, 82071, Laramie, WY, USA
6. Université Lyon 1, CNRS, Laboratoire de Biométrie et Biologie Evolutive, UMR 5558, Villeurbanne, France