1. School of Science and Engineering, University of the Sunshine Coast, Maroochydore DC, Queensland 4558, Australia
2. School of Marine and Tropical Biology, James Cook University, Townsville, Queensland 4811, Australia
3. Centre for Microscopy and Microanalysis and School of Biological Sciences, The University of Queensland, St Lucia, Queensland 4072, Australia
4. The University of Oxford, Begbroke Science Park, Sandy Lane, Yarnton OX5 1PF, UK
5. Previously Queensland Department of Environment and Heritage Protection, GPO Box 2454, Brisbane, Queensland 4001, Australia