1. Division of Bioengineering and Department of Mechanical Engineering, National University of Singapore, Singapore 117576, Republic of Singapore
2. Research Centre of Excellence in Mechanobiology, National University of Singapore, Singapore 117411, Republic of Singapore
3. Department of Molecular Cell Biology, Weizmann Institute of Science, Rehovot 76100, Israel
4. Department of Biological Sciences, Columbia University, New York, NY 10027, USA