1. School of Earth, Atmospheric and Environmental Sciences, and Williamson Research Centre for Molecular Environmental Science, University of Manchester, Manchester M13 9PL, UK
2. Geomicrobiology, Center for Applied Geoscience, University of Tuebingen, Sigwartstrasse 10, 72076 Tuebingen, Germany
3. Manchester Institute of Biotechnology, University of Manchester, Manchester M1 7DN, UK
4. The Centre for Process Innovation, CPI, Wilton Centre, Wilton, Redcar TS10 4RF, UK