1. School of Biological Sciences, The University of Adelaide, North Terrace, Adelaide, South Australia 5005, Australia
2. South Australian Museum, North Terrace, Adelaide, South Australia 5001, Australia
3. School of Medicine, Medical Sciences and Nutrition, University of Aberdeen, Aberdeen AB25 2ZD, UK
4. School of Engineering and Computer Science, Medical and Biological Engineering Research Group, University of Hull, Hull HU6 7RX, UK
5. Research Department of Cell and Developmental Biology, UCL, University College London, Anatomy Building, Gower Street, London WCIE 6BT, UK