1. School of Mathematics, Cardiff University, Senghennydd Road, Cardiff CF24 4AG, UK
2. Institute for Medicine and Engineering, University of Pennsylvania, 3340 Smith Walk, Philadelphia, PA, USA
3. Departments of Physiology and Physics and Astronomy, Institute for Medicine and Engineering, University of Pennsylvania, 3340 Smith Walk, Philadelphia, PA, USA
4. Mathematical Institute, University of Oxford, Woodstock Road, Oxford OX2 6GG, UK