1. Department of Entomology and Nematology, University of California-Davis, 1282 Academic Surge, Davis, CA 95616, USA
2. Department of Ecology and Evolutionary Biology, University of Connecticut, 75 North Eagleville Road, Storrs, CT 06269-3043, USA
3. Institut für Spezielle Zoologie und Evolutionsbiologie, FSU Jena, 07743 Jena, Germany