1. Boyd Orr Centre for Population and Ecosystem Health, Institute of Biodiversity, Animal Health and Comparative Medicine, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow G12 8QQ, UK
2. Department of Bioscience, Aarhus University, Kalø, Grenåvej 14, DK-8410 Rønde, Denmark
3. College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow G12 8QQ, UK