1. Department of Biological Sciences, University of Arkansas, Fayetteville, AR 72701, USA
2. Department of Ecology and Evolutionary Biology, University of Colorado, Boulder 80309, USA
3. National Institute of Arthritis, Musculoskeletal and Skin Diseases (NIAMS), National Institutes of Health, Bethesda, MD 20892, USA
4. Department of Biological Sciences, Emporia State University, Emporia, KS 66801, USA
5. National Research and Development Centre for Riverine and Lake Fisheries, Ministry of Agriculture and Forests, Royal Government of Bhutan, Haa, Bhutan