1. Committee on Evolutionary Biology, University of Chicago, 1025 E. 57th Street, Chicago, IL 60637, USA
2. Smithsonian Tropical Research Institute, Apartado 0843-03092, Balboa, Ancón, Republic of Panamá
3. Department of Evolution, Ecology, and Organismal Biology, The Ohio State University, 318 W. 12th Ave, Columbus, OH 43210, USA
4. Greifswald University, Zoological Institute and Museum, Soldmannstr. 14, 17489 Greifswald, Germany