1. 1925 Animal genetics an introduction to the science of animal breeding XVII + 420 pages. Edinburgh & London: Oliver & Boyd. (Russian edition 1929.)
2. 1927 Organic inheritance in man. (The subject matter of a course of lectures given by the author as the first William Withering lecturer in the University of Birmingham.) XXVIII + 214 pages. Edinburgh & London: Oliver & Boyd. The genetics of sexuality in animals x+ 188 pages. Cambridge University Press.
3. 1928 Heredity Benn's Sixpenny Library no. 110 79 pages. London: Ernest Benn.
4. 1931 (With J. N. P ickard) The scientific aspects of rabbit breeding VIII + 122 pages 12 plates. Idle Bradford: Watmough.
5. 1932 An introduction to the study of sex. (160 pages.) Gollancz' outline series. London: Gollancz.