1. School of Biosciences, University of Exeter, Penryn Campus, Penryn, Cornwall TR10 9FE, UK
2. Department of Biology, University of York, Wentworth Way, York YO10 5DD, UK
3. Department of Biological and Environmental Science, Nanoscience Center, University of Jyväskylä, PL 35, 40014 Jyväskylä, Finland