1. Menzies Institute for Medical Research, University of Tasmania, Hobart, Tasmania 7000, Australia
2. School of Biological Sciences, University of Tasmania, Hobart, Tasmania 7001, Australia
3. Centre for Biological Science, University of Southampton, Highfield Campus, Southampton SO17 1BJ, UK
4. Department of Primary Industries, Parks, Water and Environment, Hobart, Tasmania 7000, Australia
5. School of Medicine, University of Tasmania, Hobart, Tasmania 7000, Australia