1. Biodiversity and Macroecology Group, Department of Animal and Plant Sciences, University of Sheffield, Sheffield S10 2TN, UK
2. Department of Ecology, College of Urban and Environmental Sciences, Peking University, Beijing 100871, China
3. Department of Town and Regional Planning, University of Sheffield, Sheffield S10 2TN, UK
4. Durrell Institute for Conservation and Ecology (DICE), University of Kent, Canterbury, Kent CT2 7NR, UK