1. Environment Department, Faculty of Sciences, University of York, Heslington, York YO10 5NG, UK
2. Department of Epidemiology and Population Health, Institute of Infection and Global Health, Leahurst Campus, University of Liverpool, Neston CH64 7TE, UK
3. School of Veterinary Medicine and Science, University of Nottingham, Sutton Bonington Campus, Sutton Bonington, Leicestershire LE12 5RD, UK