1. LGL Ecological Research Associates Inc., Bryan, TX 77801, USA
2. Department of Fisheries and Wildlife, Oregon State University, Corvallis, OR 97331, USA
3. Oregon Hatchery Research Center, Alsea, OR 97324, USA
4. Geology and Geophysics, College of Earth, Ocean and Atmospheric Sciences, Oregon State University, Corvallis, OR 97331, USA
5. School of Aquatic and Fishery Sciences, University of Washington, Seattle, WA 98195, USA
6. Department of Biology, University of North Carolina, Chapel Hill, NC 27599, USA