1. Department of Biological Sciences, University of Toronto Scarborough, Toronto, Canada M1C1A4
2. Department of Ecology and Evolutionary Biology, University of Toronto Scarborough, Toronto, Canada M1C1A4
3. Blaustein Institutes for Desert Research, Ben-Gurion University of the Negev, Sede Boqer Campus, Beersheba 8499000 Israel
4. Department of Entomology, Volcani Center, Bet Dagan 50250, Israel