1. German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig, Puschstrasse 4, 04103 Leipzig, Germany
2. Leipzig University, Institute of Biology, Puschstrasse 4, 04103 Leipzig, Germany
3. J.F. Blumenbach Institute of Zoology and Anthropology, University of Goettingen, Untere Karspuele 2, Goettingen 37073, Germany
4. Department Biological Sciences, University of Calgary, Calgary, Alberta, Canada T2N 1N4