1. Department of Paleontology, San Diego Natural History Museum, San Diego, CA 92101, USA
2. University of California Museum of Paleontology, Berkeley, CA 94720, USA
3. Department of Earth Sciences, University of Oregon, Eugene, OR 97403, USA
4. Center for International Collaboration and Advanced Studies in Primatology, Kyoto University Primate Research Institute, Inuyama, Aichi 484-8506, Japan
5. Center for the Evolutionary Origins of Human Behavior, Kyoto University, Inuyama, Aichi 484-8506, Japan
6. Negaunee Integrative Research Center, Field Museum of Natural History, Chicago, IL 60605, USA