1. Biology Department, Woods Hole Oceanographic Institution, Woods Hole, MA 02543, USA
2. Department of Earth, Atmospheric and Planetary Science, Massachusetts Institute of Technology, Cambridge, MA 02139, USA
3. School of Environmental Sciences, University of Liverpool, Nicholson Building, Brownlow Street, Liverpool, L69 3GP, UK
4. Centre d'Etudes Biologiques de Chize, CNRS-La Rochelle University UMR7372, 79360 Villiers en Bois, France