1. Department of Biology, University of Nevada, Reno, NV 89557, USA
2. Ecology, Evolution, and Conservation Biology Interdisciplinary Program, University of Nevada, Reno, NV 89557, USA
3. Department of Biological Sciences, Fordham University, New York, NY 10458, USA
4. United States Department of Agriculture, Agricultural Research Service, Reno, NV 89512, USA
5. United States Fish and Wildlife Service, Lahontan National Fish Hatchery Complex, 1340 Financial Blvd, Suite 234, Reno, NV 89502, USA