1. Department of Integrative Biology, Michigan State University, 288 Farm Lane, Room 203, East Lansing, MI 48824-1115, USA
2. Ecology, Evolutionary Biology and Behavior, Michigan State University, 103 Giltner Hall, East Lansing, MI 48824, USA
3. Department of Zoology and Physiology, University of Wyoming, 1000 E. University Ave, Biological Science Building, Laramie, WY 82071, USA
4. Program in Ecology, Berry Center, University of Wyoming, Laramie, WY 82071, USA