1. Department of Zoology, University of TorontoToronto, Ontario M5S 3G5, Canada
2. Centre for Biodiversity and Conservation Biology, Royal Ontario MuseumToronto, Ontario M5S 2C6, Canada
3. Department of Biology, Queen's UniversityKingston Ontario K7L 3N6, Canada
4. Department of Mathematics and Statistics, Queen's UniversityKingston Ontario K7L 3N6, Canada