1. Behavioural Ecology Research Group, Department of Evolutionary Zoology, University of DebrecenH-4032, Debrecen, Egyetem tér 1., Hungary
2. Department of Mathematics, University of Bristol, University WalkBristol, BS8 1TW, UK
3. School of Biological Sciences, University of BristolWoodland Road, Bristol BS8 1UG, UK
4. Animal Ecology, Department of Ecology, Lund UniversityLund, Sweden
5. Department of Theoretical Ecology, Lund UniversityEcology Building, 223 62 Lund, Sweden