1. Department of Biological Sciences, University of South CarolinaColumbia, SC 29208, USA
2. Edward Grey Institute, Department of Zoology, University of OxfordOxford OX1 3PS, UK
3. School of Zoology, University of TasmaniaHobart, Tasmania 7001, Australia
4. Department of Ecology and Evolutionary Biology, University of ArizonaTucson, AZ 85721, USA