1. Institute of Evolutionary Biology, School of Biological Sciences, The University of Edinburgh, Ashworth LaboratoriesThe King's Buildings, Edinburgh EH9 3JT, UK
2. CSIRO EntomologyGPO Box 1700, Canberra ACT 2601, Australia
3. Division of Pathway Medicine and Centre for Infectious Diseases, University of Edinburgh Medical SchoolChancellor's Building, 49 Little France Crescent, Edinburgh EH16 4SB, UK